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The Wolves in Nature Project


Ripple, W.J. and Beschta, R.L. (2006). Linking a cougar decline, trophic cascade, and catastrophic regime shift in Zion National Park. Biological Conservation 133: 397-408.

Larsen, T. and Ripple, W.J. (2006). Modeling Gray Wolf (Canis lupus) Habitat in the Pacific Northwest, U.S.A. Journal of Conservation Planning 2(1): 30-61.

Beschta, R.L. and Ripple, W.J. (2006). River channel dynamics following extirpation of wolves in northwestern Yellowstone National Park,USA. Earth Surface Processes and Landforms 31: 1525-1539.

Ripple, W.J. and Beschta, R.L. (2006). Linking wolves to willows via risk-sensitive foraging by ungulates in the northern Yellowstone ecosystem. Forest Ecology and Management 230: 96-106.

Beschta, R.L. (2005). Reduced Cottonwood Recruitment Following Extirpation of Wolves in Yellowstone's Northern Range. Ecology 86(2): 391-403.

Beschta, R.L. and W.J. Ripple. (2005). Rapid Assessment of Riparian Cottonwood Recruitment: Middle Fork John Day River, Northeastern Oregon. Ecological Restoration. 23(3):150-156.

Ripple, W.J. and Beschta, R.L. (2005). Linking Wolves and Plants: Aldo Leopold on Trophic Cascades. BioScience July 2005/Vol. 55 No. 7: 613-621 - Abstract

Ripple, W.J. and Beschta, R.L. (2005). Refugia from browsing as reference sites for restoration planning. Western North American Naturalist 65(2): 269-273.

Ripple, W.J. and Beschta, R.L. (2005). Willow thickets protect young aspen from elk browsing after wolf reintroduction. Western North American Naturalist 65(1): 118-122.

Laliberte, Andrea S. and Ripple, William J. Range Contractions of North American Carnivores and Ungulates. BioScience. February 2004/Vol. 54 No. 2: 123-138.

Ripple W.J., Beschta R.L. 2004. Wolves and the ecology of fear: Can predation risk structure ecosystems? BioScience. 54:755-766. - Abstract

Laliberte, A.S. and Ripple W.J. (2004). Range Contractions of North American Carnivores and Ungulates - Abstract

Mystery in Yellowstone: wolves, wapiti, and the case of the disappearing aspen. Notable Notes, Oregon State University, 2004.

Ripple, William J. and Beschta, Robert L. (2004). Wolves and the Ecology of Fear: Can Predation Risk Structure Ecosystems? BioScience Augutst 2004/Vo.l 54 No. 8: 755-766.

Ripple, W.J. and Beschta, R.L. (2004). Wolves, elk, willows, and trophic cascades in the upper Gallatin Range of Southwestern Montana, USA. Forest Ecology and Management 200: 161-181.

Beschta, R.L. (2003). Cottonwoods, elk, and wolves in the Lamar Valley of Yellowstone National Park. Ecological Applications 13(5): 1295-1309.

Laliberte, Andrea S. and Ripple, William J. (2003). Wildlife Encounters by Lewis and Clark: A Spatial Analysis of Interactions between Native Americans and Wildlife. BioScience October 2003/Vol. 53 No. 10: 994-1003.

Larsen, E.J. and W.J. Ripple. (2003). Aspen age structure in the northern Yellowstone Ecosystem:USA. Forest Ecology and Management. 179: 469-482.

Powell, D. Reintroduction of Wolves May End Tree and Shrub Decline in Yellowstone. The Forestry Source (December 2003).

Ripple, W.J. and Beschta, R.L. (2003). Wolf reintroduction, predation risk, and cottonwood recovery in Yellowstone National Park. Forest Ecology and Management 184: 299-313.

Ecological Dynamics on Yellowstone's Northern Range, The National Acadamy of Science, 2002.

Humans, wolves, elk, aspen and willow, and now beetles (HWEAW+B) science workshop proceedings, February 5 & 6, 2002, Banff, Alberta.

Ripple, W. J., E. J. Larsen, R. A. Renkin, D. W. Smith. (2001). Trophic Cascades among wolves, elk and aspen on Yellowstone National Park’s northern range. Biological Conservation. 102:227-234.

Bergdahl, B. Trophic dynamics in the ecosystem in regard to the presence of wolves. Kenyon College, 2000.

Ripple, W.J. and Larsen, E.J. (2000). Historic aspen recruitment, elk, and wolves in northern Yellowstone National Park, USA. Biological Conservation 95: 361-370.

The Wolf, the Moose, and the Fir Tree: a case study of trophic interactions - Gary M. Fortier, Dept. of Small Animal Science, Delaware Valley College, 1998.